Emperor Penguins from Antarctica each express their true love with a special heartsong of their own that expresses their very being. However, the misfit Mumble cannot sing. Instead he has an extraordinary talent to tap dance with almost magical energy and expression. Nevertheless, the leadership of the colony fearfully blames the young penguin's unorthodox ways for the lean fishing that threatens them all. Defiant in the face of unjust rejection,...

When Mumble's son Erik wants to be a singer instead of a dancer like his father, they have a falling out until a problem that threatens the entire penguin colony brings them together.

In the Antarctic, every March since the beginning of time, the quest begins to find the perfect mate and start a family. This courtship will begin with a long journey - a journey that will take them hundreds of miles across the continent by foot, one by one in a single file. They will endure freezing temperatures, in brittle, icy winds and through deep, treacherous waters. They will risk starvation and attack by dangerous predators, under the harshest...